Most papers are open access and free to download (just click on the paper title to get redirected to the PDF). If you cannot find a specific article, please do get in touch with me. Equal contributions are denoted by *, empirical papers by (e) and theoretical papers by (t).

Journals – Submitted/Under review/Preprints

66(t). Anichini, M., de Heer Kloots, M., Ravignani, A. (submitted) Animal vocal rhythms in the wild, in the lab, and in silico.

65(e). Geambasu, A., Toron, L., Ravignani, A., & Levelt, C. C. (submitted). Rhythmic recursion? Human sensitivity to a Lindenmayer grammar with self-similar structure in a musical task.

64(t). Ravignani, A., Kotz, S. A. (2017, preprint). Evolution of temporal interaction: A comparative approach to social timing. PeerJ Preprints 5:e3275v1 doi: 10.7287/peerj.preprints.3275v1

63(e/t). Shillcock, R., Millidge, B., Ravignani, A. (2019, preprint). Vocal imitation can create acoustic attractors to guide mothers to pups in a crowded colony of Mexican free-tailed bats: A case-study of computational modelling in behavioural biology. PsyArXiv.

Journals – Published/In press

2020/in press

62(t). Ravignani, A., de Boer, B. (in press) Joint origins of speech and music: Testing evolutionary hypotheses on modern humans. Semiotica.

61(t). Jacoby, N., Margulis, E., Clayton, M., Hannon, E., Honing, H. Iversen, J., Klein, T.R., Mehr, S. A., Pearson, L. Peretz, I., Perlman, M., Polak, R., Ravignani, A., Savage, P.E., Steingo, G., Stevens, C., Tomlinson, G., Trainor, L., Trehub, S., Veal M., & Wald-Fuhrmann, M. (2020) Cross-Cultural Research in Music Cognition: Challenges, Insights and Recommendations. [pdfMusic Perception.

60(e/t). de Boer, B., Thompson, B., Ravignani, A., Boeckx, C. (2020). Evolutionary Dynamics Do Not Motivate a Single-Mutant Theory of Human Language. Scientific Reports.


59(t). Ravignani, A. (2019) Seeking shared ground in space. [pdfScience.

58(t). Larsson, M., Richter, J., Ravignani, A. (2019) Bipedal steps in the development of rhythmic behaviour in humans. Music & Science.

57(t). Ravignani, A. (2019) Everything you always wanted to know about sexual selection in 129 pagesJournal of Mammalogy.

56(e). Lameira, A., Eerola, T., Ravignani, A., (2019) Coupled whole-body rhythmic entrainment between two chimpanzees. Scientific Reports.

55(t). Ravignani, A., Verga, L., Greenfield, M. (2019) Interactive rhythms across species: The evolutionary biology of animal chorusing and turn-taking. Annals of the New York Academy of Sciences.

54(t). Ravignani, A. (2019) Singing seals imitate human speech. Journal of Experimental Biology.

53(t). Ravignani, A. (2019) Animal Beauty: On the evolution of biological aesthetics. Animal Behaviour.

52(t). Ravignani, A. (2019) Understanding mammals, hands-onJournal of Mammalogy.

51(e). Versace, E., Rogge, J. R., Shelton-May, N., Ravignani, A. (2019) Positional encoding in cotton-top tamarins (Saguinus oedipus). Animal Cognition.

50(t). Ravignani, A.,  Dalla Bella, S., Falk, S., Kello, C.,  Noriega, F. & Kotz, S. A. (2019) Rhythm in speech and animal vocalizations: a cross-species perspective. Annals of the New York Academy of Sciences.

49(t). Ravignani, A. & Gamba, M. (2019) Evolving Musicality. Trends in Ecology and Evolution.

48(t). Ravignani, A. (2019) Humans and other musical animals. Current Biology.

47(e). Ravignani, A.*,  Filippi, P.* & Fitch, T. (2019) Perceptual tuning and rule abstraction: insights for comparative experiments on pattern perception. i-Perception.

46(e/t). Ravignani, A. & de Reus, K. (2019). Modelling animal interactive rhythms in communicationEvolutionary Bioinformatics.

45(t). Ravignani, A. (2019). Rhythm and synchrony in animal movement and communication. Current Zoology.

44(e). Reber, S., Šlipogor, V., Oh, J., Ravignani, A., Hoeschele, M., Bugnyar, T., Fitch, W.T. (2019). Common marmosets are sensitive to simple dependencies at variable distances in an artificial grammar. Evolution & Human Behaviour.

43(t). Ravignani, A. (2019). The origins of musicality. Perception.


42(e). Ravignani, A. (2018). Timing of antisynchronous calling: A case study in a harbor seal pup (Phoca vitulina). Journal of Comparative Psychology. [Preprint article]

41(t). Ravignani, A., Thompson, B., Lumaca, M., Grube, M. (2018) Why do durations in musical rhythms conform to small integer ratios? Frontiers in Computational Neuroscience.

40(t). Kotz, S.*, Ravignani, A.*, Fitch, W.T. (2018). The evolution of rhythm processingTrends in Cognitive Sciences

39(t). Ravignani, A. (2018). Darwin, sexual selection, and the origins of musicTrends in Ecology and Evolution.

38(e). Ravignani, A., Kello, C., de Reus, K., Kotz, S. A., Dalla Bella, S., Mendez, M., Rapado, B., Rubio-Garcia, A., & de Boer, B. (2018) Ontogeny of vocal rhythms in harbour seal pups: An exploratory study. Current Zoology.

37(t). Wilson, B., Spierings, M., Ravignani, A., Mueller, J., Mintz, T., Wijnen, F., van der Kant, A., Smith., K., Rey, A. (2018) Non-adjacent dependency learning in humans and other animalsTopics in Cognitive Sciences.

36(t). Lumaca, M., Ravignani, A., Baggio, G. (2018) Music evolution in the laboratory: Cultural transmission meets neurophysiologyFrontiers in Neuroscience.

35(t). Ravignani, A., Thompson, B., Filippi, P. (2018). The evolution of musicality: What can be learned from language evolution research? Frontiers in Neuroscience.

34(t). Ravignani, A. (2018). Comment on “Temporal and spatial variation in harbor seal (Phoca vitulina L.) roar calls from southern Scandinavia” [J. Acoust. Soc. Am. 141, 1824-1834 (2017)]. The Journal of the Acoustical Society of America. 143(1):504-508. [Author’s PDF, (C) 2018 Acoustical Society of America]

33(e). Ravignani, A.*, Thompson, B.*, Grossi, T., Delgado, T. & Kirby, S. (2018) Evolving building blocks of rhythm: How human cognition creates music via cultural transmission Annals of the New York Academy of Sciences (preprint here).

32(e). Ravignani, A. (2018) Spontaneous rhythms in a harbor seal pup calls. BMC Research Notes 11(3).

31(t). Ravignani, A., Verhoef, T. (2018, preprint available). Which melodic universals emerge from repeated signaling games? Artificial Life. 24 (2).


30(t). Ravignani, A., Madison, G. (2017). The paradox of isochrony in the evolution of human rhythm. Frontiers in Psychology. 

29(t). Ravignani, A., Honing H., & Kotz S.A. (2017) Editorial: The Evolution of Rhythm Cognition: Timing in Music and Speech. Frontiers in Human Neuroscience 11:303.

28(t). Ravignani, A. (2017) Interdisciplinary debate: Agree on definitions of synchrony. Nature. 545, 158.

27(t). Ravignani, A., Thompson, B. (2017) A note on ‘Noam Chomsky – What kind of creatures are we?’ Language in Society. 46(3), 446-447.

26(e). Ravignani, A.*, Gross, S.*, Garcia, M., Rubio-Garcia, A., & de Boer, B. (2017) How small could a pup sound? The physical bases of signalling body size in harbour seals. Current Zoology. doi:10.1093/cz/zox026.

25(e/t). Ravignani, A. (2017) Visualizing and interpreting rhythmic patterns using phase space plots. Music Perception. 34(5), 557-568.

24(t). Ravignani, A.* & Norton, P.* (2017) Measuring rhythmic complexity: A primer to quantify and compare temporal structure in speech, movement, and animal vocalizations. Journal of Language Evolution. 2 (1), 4-19.

23(e). Ravignani, A.* & Sonnweber, R.* (2017) Chimpanzees process structural isomorphisms across sensory modalities. Cognition. 161, 74-79.


22(e). Ravignani, A., Delgado, T. & Kirby, S. (2016) Musical evolution in the lab exhibits rhythmic universals. Nature Human Behaviour. 1:1.

21(e). Jadoul, Y.*, Ravignani, A.*, Thompson, B.*, Filippi, P., & de Boer, B. (2016). Seeking temporal predictability in speech: Comparing statistical approaches on 18 world languages. Frontiers in Human Neuroscience. 10:586.

20(t). Ravignani, A. & Cook, P. (2016) The evolutionary biology of dance, without frills. Current Biology. 26:19.

19(e). Geambasu, A.*, Ravignani, A.*, & Levelt, C. C. (2016). Preliminary experiments on human sensitivity to rhythmic structure in a grammar with recursive self-similarity. Frontiers in Neuroscience. 10:281.

18(t). Ravignani, A., Fitch, W. T., Hanke, F. D., Heinrich, T., Hurgitsch, B., Kotz, S. A., Scharff, C., Stoeger, A.S., & de Boer, B. (2016). What pinnipeds have to say about human speech, music, and the evolution of rhythm. Frontiers in Neuroscience. 10:274.


17(t). Ravignani A. (2015). Evolving perceptual biases for antisynchrony: A form of temporal coordination beyond synchrony. Frontiers in Neuroscience. 9:339.

16(e). Ravignani, A., Westphal-Fitch, G., Aust, U., Schlumpp, M. & Fitch, T. (2015) More than one way to see it: Individual heuristics in avian visual computation. Cognition.

15(e). Sonnweber, R., Ravignani, A., Stobbe, N., Schiestl, G., Wallner, B. & Fitch, T. (2015) Rank-dependent grooming patterns and cortisol alleviation in Barbary macaques. American Journal of Primatology.

14(t). Ravignani, A.* & Sonnweber, R.* (2015) Measuring teaching through hormones and time series analysis: Towards a comparative framework. Behavioral and Brain Sciences.

13(e). Sonnweber R-S, Ravignani A, Fitch WT. (2015) Non-adjacent visual dependency learning in chimpanzees. Animal Cognition.


12(t). Ravignani, A., Bowling, D. & Fitch, T. (2014) Chorusing, synchrony and the evolutionary functions of rhythmFrontiers in Psychology, 5:1118.

11(e). Fuhrmann, D., Ravignani, A., Marshall-Pescini, S. & Whiten, A. (2014). Synchrony and motor mimicking in chimpanzee observational learningScientific Reports, 4: 5283.

10(t). Ravignani, A. (2014) Chronometry for the chorusing herd: Hamilton’s legacy on context-dependent acoustic signalling—a comment on Herbers (2013). Biology letters10(1), 20131018.

9(t). Ravignani, A.*, Martins, M.* & Fitch, W.T. (2014) Vocal learning, prosody and basal ganglia: Don’t underestimate their complexity. Behavioral and Brain Sciences.


8(e). Ravignani A, Sonnweber R-S, Stobbe N, Fitch WT. (2013) Action at a distance: dependency sensitivity in a New World primate. Biology Letters, 9: 20130852.

7(e). Ravignani, A.; Olivera, V.M.; Gingras, B.; Hofer, R.; Hernandez, C.R.; Sonnweber, R.-S.; Fitch, W.T. (2013) Primate Drum Kit: A System for Studying Acoustic Pattern Production by Non-Human Primates Using Acceleration and Strain Sensors. Sensors, 13, 9790-9820.

Invited Papers in Popular Science Journals

6(t). Ravignani, A., Chiandetti, C. & Gamba, M. (2018) L’evoluzione del ritmo. Le Scienze. [The evolution of rhythm. Scientific American, Italian edition.]

Full Papers in Peer-reviewed Proceedings

5(t). Ravignani, A., Bowling, D. & Kirby, S. (2014) The psychology of biological clocks: a new framework for the evolution of rhythm. The Evolution of Language: Proceedings of the 10th international conference. (pp. 262-269) ISBN 978-981-4603-62-1.

4(t). Ravignani, A.; Gingras, B.; Asano, R.; Sonnweber, R.; Matellan, V.; Fitch, W.T. (2013) The Evolution of Rhythmic Cognition: New Perspectives and Technologies in Comparative Research. In Proceedings of the 35th Annual Conference of the Cognitive Science Society; Knauff, M., Pauen M., Sebanz, N. & Wachsmuth, I., Eds.; Cognitive Science Society: Austin, TX, USA; pp. 1199-1204.

3(e). Ravignani, A. & Fitch, W.T. (2012) Sonification of experimental parameters as a new method for efficient coding of behavior. Proceedings of Measuring Behavior 2012, 8th International Conference on Methods and Techniques in Behavioral Research. Edited by A.J. Spink, F. Grieco, O.E. Krips, L.W.S. Loijens, L.P.J.J. Noldus, and P.H. Zimmerman ISBN 978-90-74821-87-2 (pg.376- 379).


2(t). Ravignani, A., Chiandetti, C., Kotz, S. (2018) Rhythm and music in animal signals. Encyclopedia of Animal Behavior, Second Edition (Elsevier).

1(t). Martins, M., Raju, A. & Ravignani, A. (2014) Evaluating the role of quantitative modeling in language evolution. The Past, Present and Future of Language Evolution Research. (pp.84-93) ISBN 978-4-9906340-1-8.

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